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This work was funded by the Wellcome Trust (WT098051). G.S.V. is funded by a Cancer Research UK Senior Cancer Fellowship (C22324/A23015) and work in his lab is also funded by the European Research Council, Kay Kendall Leukaemia Fund, Blood Cancer UK, Leukemia Lymphoma Society and Rise Tide Foundation for Clinical Cancer Research. M.A.F. is funded by a Wellcome Clinical Fellowship.